>Thanks Michael. What do you think of my theory that she started laying drones and the bees killed her?
Likely.
>So the queen is definitely laying drones - but how is there a queen cell?
http://www.bushfarms.com/huber.htm#maleeggsinroyalcells"It is a singular fact, that the females, whose fecundation has been retarded, sometimes lay the eggs of males in royal cells. I shall prove, in the history of swarms, that immediately when queens, in the natural state, begin their great laying of male eggs, the workers construct numerous royal cells. Undoubtedly, there is some secret relation between the appearance of male eggs and the construction of these cells; for it is a law of nature from which bees never derogate. It is not surprising, therefore, that such cells are constructed in hives governed by queens laying the eggs of males only. It is no longer extraordinary that these queens deposit in the royal cells, eggs of the only species they can lay, for in general their instinct seems affected. But what I cannot comprehend is, why the bees take exactly the same care of the male eggs deposited in royal cells, as of those that should become queens. They provide them more plentifully with food, they build up the cells as if containing a royal worm; in a word, they labour with such regularity that we have frequently been deceived. More than once, in the firm persuasion of finding royal nymphs, we have opened the cells after they were sealed, yet the nymph of a drone always appeared. Here the instinct of the workers seemed defective. In the natural state, they can accurately distinguish the male worms from those of common bees, as they never fail giving a particular covering to the cells containing the former. Why then can they no longer distinguish the worms of drones, when deposited in the royal cell? The fact deserves much attention. I am convinced that to investigate the instinct of animals, we must carefully observe where it appears to err."--Francis Huber, New Observations on Bees Vol I Letter III, June 1787.
http://www.bushfarms.com/huber.htm#onlymaleeggs"Speaking of females laying male eggs alone, I have already expressed my surprise that bees bestow, on those deposited in royal cells, such care and attention as to feed the worms proceeding from them, and, at the period of transformation, to close them up. But I know not, Sir, why I omitted to observe that, after sealing the royal cells, the workers build them up, and sit on them until the last metamorphosis of the included male. (Translators note: It is difficult to discover whether the author thinks, as some Naturalists, that bees are instrumental in hatching the eggs. T.) The treatment of the royal cells where fertile workers lay the eggs of drones is very different. They begin indeed with bestowing every care on their eggs and worms; they close the cells at a suitable time, but never fail to destroy them three days afterwards."--Francis Huber, New Observations on Bees, Vol I Letter V, August 1788.