Fish, what Jim said, plus maybe brood chill as this was part of the original removed comb. Either way, only a small patch and rest of the hive is thriving.
So, lack of stores, as Jim text
Brood chill
Brood over heat
Only a small patch of brood is affected so I would rule out Disease or varroa.
Again, great close up pic. Only a few cells in the pic but I see no varroa frass.