well, to quote michael bush, "it depends".
in general, the bees will want 15-20% drone cells. under normal circumstances (for the bees), these cells are generally distributed around the outside of the broodnest (tops and bottom corners of brood frames, frames on the outside of the broodnest).
when you introduce foundationless frames to a colony with with nearly all worker sized cells (ie, from foundation), the full sheets of drone comb will look alarming...and if you remove each sheet of drone comb and "start over" with a new foundationless frame, you will get the same result...all drone cells.
bees will also build drone sized cells if they are building comb to store honey (less wax to store more honey).
but in the end, the bees will do their best to raise the number of drones they want. removing drone brood will only be a waste of the resources invested in them....if they do not emerge, the bees will simply start over.
another related issue is drone brood in the honey supers. comb that has had brood raised in it (and has leftover cocoons) is difficult for the bees to rework into drone comb....so, they look for comb that has not had brood raised in it and is easily worked....the honey super! the queen will generally not cross the honey barrier to lay above if there is sufficent space to raise drones in the broodnest...but the universal use of foundation prevents this from happening...hence, the queen exluder is "needed" to keep the drone brood out of the honey super.
deknow